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        <datestamp>2026-09-21T07:24:56Z</datestamp>
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          <dc:title>A new taxon of hybodontiform (Chondrichthyes, Elasmobranchii, Hybodontiformes) from the Late Triassic of Arizona, U.S.A., and its biostratigraphic importance within the Chinle Formation</dc:title>
          <dc:creator>Emily Keeble (8336253)</dc:creator>
          <dc:creator>Ben T. Kligman (6319655)</dc:creator>
          <dc:creator>Adam D. Marsh (5843300)</dc:creator>
          <dc:creator>Michelle R. Stocker (8920124)</dc:creator>
          <dc:creator>Sterling J. Nesbitt (7342976)</dc:creator>
          <dc:subject>Molecular Biology</dc:subject>
          <dc:subject>Evolutionary Biology</dc:subject>
          <dc:subject>Environmental Sciences not elsewhere classified</dc:subject>
          <dc:subject>Chemical Sciences not elsewhere classified</dc:subject>
          <dc:subject>Ecology</dc:subject>
          <dc:subject>Biological Sciences not elsewhere classified</dc:subject>
          <dc:subject>Inorganic Chemistry</dc:subject>
          <dc:subject>small lateral cusps</dc:subject>
          <dc:subject>productive microvertebrate site</dc:subject>
          <dc:subject>principal (= center</dc:subject>
          <dc:subject>lack ornamentation apart</dc:subject>
          <dc:subject>biostratigraphic importance within</dc:subject>
          <dc:subject>lonchidion minimorsus &lt;/</dc:subject>
          <dc:subject>lonchidion &lt;/</dc:subject>
          <dc:subject>biostratigraphic correlation</dc:subject>
          <dc:subject>present within</dc:subject>
          <dc:subject>localities within</dc:subject>
          <dc:subject>wide distributions</dc:subject>
          <dc:subject>upper triassic</dc:subject>
          <dc:subject>taxonomy difficult</dc:subject>
          <dc:subject>stratigraphic sequence</dc:subject>
          <dc:subject>stratigraphic gap</dc:subject>
          <dc:subject>sonsela member</dc:subject>
          <dc:subject>recognized distribution</dc:subject>
          <dc:subject>possesses teeth</dc:subject>
          <dc:subject>one population</dc:subject>
          <dc:subject>new taxon</dc:subject>
          <dc:subject>mesiodistally twice</dc:subject>
          <dc:subject>mainly relating</dc:subject>
          <dc:subject>late triassic</dc:subject>
          <dc:subject>labiolingually wide</dc:subject>
          <dc:subject>labial peg</dc:subject>
          <dc:subject>increases diversity</dc:subject>
          <dc:subject>higher levels</dc:subject>
          <dc:subject>high variation</dc:subject>
          <dc:subject>excellent candidates</dc:subject>
          <dc:subject>dockum group</dc:subject>
          <dc:subject>distinctive dentitions</dc:subject>
          <dc:subject>distal edge</dc:subject>
          <dc:subject>common occurrence</dc:subject>
          <dc:subject>chinle formation</dc:subject>
          <dc:subject>appears useful</dc:subject>
          <dc:subject>anatomical variation</dc:subject>
          <dc:description>&lt;p&gt;Elasmobranchs are excellent candidates for biostratigraphy, with distinctive dentitions and wide distributions over a long period of time. However, the high variation in tooth shapes present within some groups of chondrichthyans makes disentangling species variation from higher levels of taxonomy difficult. Here, we describe a sample of over 250 isolated tooth crowns of a new taxon of hybodontiform from a productive microvertebrate site from the Sonsela Member of the Chinle Formation, Arizona, U.S.A. This sample is assigned to a new taxon, &lt;i&gt;Lonchidion minimorsus&lt;/i&gt; sp. nov., which possesses teeth that are mesiodistally twice as long as they are labiolingually wide, include no or small lateral cusps, have a low to moderately defined labial peg consistent with other members of &lt;i&gt;Lonchidion&lt;/i&gt;, and lack ornamentation apart from the common occurrence of two or three lingually oriented cristae. Anatomical variation is present within the sample, mainly relating to the presence and/or size of the cusplets and labial peg, number of cristae, and the angle of the distal edge of the principal (= center) cusp. Three-dimensional geometric morphometrics show variation in one population and we hope that in the future further populations or species can be compared. &lt;i&gt;Lonchidion minimorsus&lt;/i&gt; sp. nov. increases diversity of the genus &lt;i&gt;Lonchidion&lt;/i&gt; and fills a stratigraphic gap in the recognized distribution of chondrichthyans from the Upper Triassic of the western U.S.A. Together, the stratigraphic sequence of &lt;i&gt;Lonchidion&lt;/i&gt; appears useful for biostratigraphic correlation of localities within and across the Chinle Formation and Dockum Group in the southwestern U.S.A.&lt;/p&gt;</dc:description>
          <dc:date>2026-09-21T07:24:56Z</dc:date>
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          <dc:identifier>10.6084/m9.figshare.33950845.v1</dc:identifier>
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          <dc:rights>CC BY 4.0</dc:rights>
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